Betty frantically tries to keep Daniel focused on work as he reverts to his old hard-partying ways, including romancing a multitude of stunning models. Things go from bad to worse when a tell-all writer (Emmy winner Leslie Jordan, "Will & Grace," "Boston Legal") in possession of Fey Sommer's diaries starts stalking the Meades. Another Suarez, Ignacio, feels the heat when Immigration caseworker Constance (guest star Octavia Spencer) pressures him to make their relationship a romantic one, and Wilhelmina and Alexis both try to get their hands on Fey’s diaries -- but for different reasons.
Betty, fed up with Daniel, Christina and Henry and how they've all disappointed her, does her best to avoid all three. Meanwhile, love's all around at Mode, as Alexis gets asked out on her first date as a woman; Amanda begins a flirtation with a designer (guest star Mykel Shannon Jenkins) who she assumes is gay; Wilhelmina struggles to deal with the tireless advances of her latest conquest, and Hilda gets a surprise from Santos (guest star Kevin Alejandro). Meanwhile, Daniel goes it alone in dealing with the aftermath of his one-time hook-up with Petra.
Betty struggles to come up with the cash to solve the latest Suarez family dilemma, all while planning Mode's big assistants week bash at a medieval themed restaurant, a place with an unlikely link to Amanda's past. Meanwhile, Daniel tries to heed Betty's advice about his sexual addiction issues; Alexis considers Rodrigo's offer (guest star Cristian de la Fuente); Hilda faces a tough cosmetology school exam, and Wilhelmina becomes even bolder in her plans for total Mode domination.
Betty and her family go to Guadalajara to solve Ignacio's immigration problems, and Betty questions her Aunt Mirta (Oscar, Emmy, Tony and Grammy Award winner Rita Moreno) and cousin Clara about her late mother. Meanwhile, Alexis gets a visit from Jordan (guest star Rebecca Gayheart), the woman she dated before her "death" and operation; Marc is jealous when Amanda seemingly has a new best gay friend in Tavares, and Wilhelmina gives the new man in her life a makeover.
Betty and Henry's romance heats up. Meanwhile, Wilhelmina makes great strides in her ambitious power play, as Daniel descends further in a downward spiral. The Suarez family trip to Mexico takes a dangerous turn, with old ghosts creating new problems. Explosive secrets will be revealed, and everything will be tied together by a beloved Broadway musical. Kristin Chenoweth, Angélica Vale and Gina Gershon guest star.
Betty's life is in chaos after a series of very unfortunate events, but despite all that's happened, she's still in denial about one thing -- how much Henry's departure has affected her. Meanwhile, Amanda turns to junk food and to Marc in dealing with the revelation that Fey Sommers was her real mother, and Wilhelmina schemes to turn the tragic Meade family events to her advantage.
Betty doesn't know what to do when she finds out Wilhelmina's hunky new bodyguard is taking the job to a whole new level.
With Henry back in her life (sort of), Betty finds herself distracted by yet another guy - Gio (Freddy Rodriguez), Mode's new sandwich guy, who both annoys and intrigues her.
Betty takes steps toward becoming a writer and invests in a class that proves to be so challenging, she tosses her own writing assignment and instead submits a Mode story written by a woman who survived a bear attack -- with disastrous results.
Betty tries Internet dating and gets asked out on a bowling date; Marc finds himself attracted to someone who is not his type; Danny and Alexis ready themselves for dinner with stepmom-to-be Wilhelmina; Justin ditches school.
Betty and Henry continue their illicit romance and plan a secret date to "Wicked," but things don't go as planned when Daniel, thinking she's dating Gio (Freddy Rodriguez), gifts her with tickets for the two of them instead.
America Ferrera
Betty Suarez
Eric Mabius
Daniel Meade
Tony Plana
Ignacio Suarez
Ana Ortiz
Hilda Suarez
Becki Newton
Amanda Tanen